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      nutrition exercised by the other parts are ancillary to the activity

      of the heart. It is the part of the dominating organ to achieve the

      final result, as of the physician's efforts to be directed towards

      health, and not to be occupied with subordinate offices.

      Certainly, however, all saguineous animals have the supreme organ of

      the sensefaculties in the heart, for it is here that we must look

      for the common sensorium belonging to all the sense-organs. These in

      two cases, taste and touch, can be clearly seen to extend to the

      heart, and hence the others also must lead to it, for in it the

      other organs may possibly initiate changes, whereas with the upper

      region of the body taste and touch have no connexion. Apart from these

      considerations, if the life is always located in this part,

      evidently the principle of sensation must be situated there too, for

      it is qua animal that an animal is said to be a living thing, and it

      is called animal because endowed with sensation. Elsewhere in other

      works we have stated the reasons why some of the sense-organs are,

      as is evident, connected with the heart, while others are situated

      in the head. (It is this fact that causes some people to think that it

      is in virtue of the brain that the function of perception belongs to

      animals.)

      4

      Thus if, on the one hand, we look to the observed facts, what we

      have said makes it clear that the source of the sensitive soul,

      together with that connected with growth and nutrition, is situated in

      this organ and in the central one of the three divisions of the

      body. But it follows by deduction also; for we see that in every case,

      when several results are open to her, Nature always brings to pass the

      best. Now if both principles are located in the midst of the

      substance, the two parts of the body, viz. that which elaborates and

      that which receives the nutriment in its final form will best

      perform their appropriate function; for the soul will then be close to

      each, and the central situation which it will, as such, occupy is

      the position of a dominating power.

      Further, that which employs an instrument and the instrument it

      employs must be distinct (and must be spatially diverse too, if

      possible, as in capacity), just as the flute and that which plays

      it-the hand-are diverse. Thus if animal is defined by the possession

      of sensitive soul, this soul must in the sanguineous animals be in the

      heart, and, in the bloodless ones, in the corresponding part of

      their body. But in animals all the members and the whole body

      possess some connate warmth of constitution, and hence when alive they

      are observed to be warm, but when dead and deprived of life they are

      the opposite. Indeed, the source of this warmth must be in the heart

      in sanguineous animals, and in the case of bloodless animals in the

      corresponding organ, for, though all parts of the body by means of

      their natural heat elaborate and concoct the nutriment, the

      governing organ takes the chief share in this process. Hence, though

      the other members become cold, life remains; but when the warmth

      here is quenched, death always ensues, because the source of heat in

      all the other members depends on this, and the soul is, as it were,

      set aglow with fire in this part, which in sanguineous animals is

      the heart and in the bloodless order the analogous member. Hence, of

      necessity, life must be coincident with the maintenance of heat, and

      what we call death is its destruction.

      5

      However, it is to be noticed that there are two ways in which fire

      ceases to exist; it may go out either by exhaustion or by

      extinction. That which is self-caused we call exhaustion, that due

      to its opposites extinction. [The former is that due to old age, the

      latter to violence.] But either of these ways in which fire ceases

      to be may be brought about by the same cause, for, when there is a

      deficiency of nutriment and the warmth can obtain no maintenance,

      the fire fails; and the reason is that the opposite, checking

      digestion, prevents the fire from being fed. But in other cases the

      result is exhaustion,-when the heat accumulates excessively owing to

      lack of respiration and of refrigeration. For in this case what

      happens is that the heat, accumulating in great quantity, quickly uses

      up its nutriment and consumes it all before more is sent up by

      evaporation. Hence not only is a smaller fire readily put out by a

      large one, but of itself the candle flame is consumed when inserted in

      a large blaze just as is the case with any other combustible. The

      reason is that the nutriment in the flame is seized by the larger

      one before fresh fuel can be added, for fire is ever coming into being

      and rushing just like a river, but so speedily as to elude

      observation.

      Clearly therefore, if the bodily heat must be conserved (as is

      necessary if life is to continue), there must be some way of cooling

      the heat resident in the source of warmth. Take as an illustration

      what occurs when coals are confined in a brazier. If they are kept

      covered up continuously by the so-called 'choker', they are quickly

      extinguished, but, if the lid is in rapid alternation lifted up and

      put on again they remain glowing for a long time. Banking up a fire

      also keeps it in, for the ashes, being porous, do not prevent the

      passage of air, and again they enable it to resist extinction by the

      surrounding air by means of the supply of heat which it possesses.

      However, we have stated in The Problems the reasons why these

      operations, namely banking up and covering up a fire, have the

      opposite effects (in the one case the fire goes out, in the other it

      continues alive for a considerable time).

      6

      Everything living has soul, and it, as we have said, cannot exist

      without the presence of heat in the constitution. In plants the

      natural heat is sufficiently well kept alive by the aid which their

      nutriment and the surrounding air supply. For the food has a cooling

      effect [as it enters, just as it has in man] when first it is taken

      in, whereas abstinence from food produces heat and thirst. The air, if

      it be motionless, becomes hot, but by the entry of food a motion is

      set up which lasts until digestion is completed and so cools it. If

      the surrounding air is excessively cold owing to the time of year,

      there being severe frost, plants shrivel, or if, in the extreme

      heats of summer the moisture drawn from the ground cannot produce

      its cooling effect, the heat comes to an end by exhaustion. Trees

      suffering at such seasons are said to be blighted or star-stricken.

      Hence the practice of laying beneath the roots stones of certain

      species or water in pots, for the purpose of cooling the roots of

      the plants.

      Some animals pass their life in the water, others in the air, and

      therefore these media furnish the source and means of refrigeration,

      water in the one case, air in the other. We must proceed-and it will

      require further application on our part-to give an account of the
    r />   way and manner in which this refrigeration occurs.

      7

      A few of the previous physical philosophers have spoken of

      respiration. The reason, however, why it exists in animals they have

      either not declared or, when they have, their statements are not

      correct and show a comparative lack of acquaintance with the facts.

      Moreover they assert that all animals respire-which is untrue. Hence

      these points must first claim our attention, in order that we may

      not be thought to make unsubstantiated charges against authors no

      longer alive.

      First then, it is evident that all animals with lungs breathe, but

      in some cases breathing animals have a bloodless and spongy lung,

      and then there is less need for respiration. These animals can

      remain under water for a time, which relatively to their bodily

      strength, is considerable. All oviparous animals, e.g. the frog-tribe,

      have a spongy lung. Also hemydes and tortoises can remain for a long

      time immersed in water; for their lung, containing little blood, has

      not much heat. Hence, when once it is inflated, it itself, by means of

      its motion, produces a cooling effect and enables the animal to remain

      immersed for a long time. Suffocation, however, always ensues if the

      animal is forced to hold its breath for too long a time, for none of

      this class take in water in the way fishes do. On the other hand,

      animals which have the lung charged with blood have greater need of

      respiration on account of the amount of their heat, while none at

      all of the others which do not possess lungs breathe.

      8

      Democritus of Abdera and certain others who have treated of

      respiration, while saying nothing definite about the lungless animals,

      nevertheless seem to speak as if all breathed. But Anaxagoras and

      Diogenes both maintain that all breathe, and state the manner in which

      fishes and oysters respire. Anaxagoras says that when fishes discharge

      water through their gills, air is formed in the mouth, for there can

      be no vacuum, and that it is by drawing in this that they respire.

      Diogenes' statement is that, when they discharge water through their

      gills, they suck the air out of the water surrounding the mouth by

      means of the vacuum formed in the mouth, for he believes there is

      air in the water.

      But these theories are untenable. Firstly, they state only what is

      the common element in both operations and so leave out the half of the

      matter. For what goes by the name of respiration consists, on the

      one hand, of inhalation, and, on the other, of the exhalation of

      breath; but, about the latter they say nothing, nor do they describe

      how such animals emit their breath. Indeed, explanation is for them

      impossible for, when the creatures respire, they must discharge

      their breath by the same passage as that by which they draw it in, and

      this must happen in alternation. Hence, as a result, they must take

      the water into their mouth at the same time as they breathe out. But

      the air and the water must meet and obstruct each other. Further, when

      they discharge the water they must emit their breath by the mouth or

      the gills, and the result will be that they will breathe in and

      breathe out at the same time, for it is at that moment that

      respiration is said to occur. But it is impossible that they should do

      both at the same time. Hence, if respiring creatures must both

      exhale and inhale the air, and if none of these animals can breathe

      out, evidently none can respire at all.

      9

      Further, the assertion that they draw in air out of the mouth or out

      of the water by means of the mouth is an impossibility, for, not

      having a lung, they have no windpipe; rather the stomach is closely

      juxtaposed to the mouth, so that they must do the sucking with the

      stomach. But in that case the other animals would do so also, which is

      not the truth; and the water-animals also would be seen to do it

      when out of the water, whereas quite evidently they do not. Further,

      in all animals that respire and draw breath there is to be observed

      a certain motion in the part of the body which draws in the air, but

      in the fishes this does not occur. Fishes do not appear to move any of

      the parts in the region of the stomach, except the gills alone, and

      these move both when they are in the water and when they are thrown on

      to dry land and gasp. Moreover, always when respiring animals are

      killed by being suffocated in water, bubbles are formed of the air

      which is forcibly discharged, as happens, e.g. when one forces a

      tortoise or a frog or any other animal of a similar class to stay

      beneath water. But with fishes this result never occurs, in whatsoever

      way we try to obtain it, since they do not contain air drawn from an

      external source. Again, the manner of respiration said to exist in

      them might occur in the case of men also when they are under water.

      For if fishes draw in air out of the surrounding water by means of

      their mouth why should not men too and other animals do so also;

      they should also, in the same way as fishes, draw in air out of the

      mouth. If in the former case it were possible, so also should it be in

      the latter. But, since in the one it is not so, neither does it

      occur in the other. Furthermore, why do fishes, if they respire, die

      in the air and gasp (as can be seen) as in suffocation? It is not want

      of food that produces this effect upon them, and the reason given by

      Diogenes is foolish, for he says that in air they take in too much air

      and hence die, but in the water they take in a moderate amount. But

      that should be a possible occurrence with land animals also; as

      facts are, however, no land animal seems to be suffocated by excessive

      respiration. Again, if all animals breathe, insects must do so also.

      many of them seem to live though divided not merely into two, but into

      several parts, e.g. the class called Scolopendra. But how can they,

      when thus divided, breathe, and what is the organ they employ? The

      main reason why these writers have not given a good account of these

      facts is that they have no acquaintance with the internal organs,

      and that they did not accept the doctrine that there is a final

      cause for whatever Nature does. If they had asked for what purpose

      respiration exists in animals, and had considered this with

      reference to the organs, e.g. the gills and the lungs, they would have

      discovered the reason more speedily.

      10

      Democritus, however, does teach that in the breathing animals

      there is a certain result produced by respiration; he asserts that

      it prevents the soul from being extruded from the body.

      Nevertheless, he by no means asserts that it is for this purpose

      that Nature so contrives it, for he, like the other physical

      philosophers, altogether fails to attain to any such explanation.

      His statement is that the soul and the hot element are identical,

      being the primary forms among the spherical particles. Hence, when

      these are being crushed together by the surrounding atmosphere

      thrusting them out, respiration, accord
    ing to his account, comes in to

      succour them. For in the air there are many of those particles which

      he calls mind and soul. Hence, when we breathe and the air enters,

      these enter along with it, and by their action cancel the pressure,

      thus preventing the expulsion of the soul which resides in the animal.

      This explains why life and death are bound up with the taking in and

      letting out of the breath; for death occurs when the compression by

      the surrounding air gains the upper hand, and, the animal being unable

      to respire, the air from outside can no longer enter and counteract

      the compression. Death is the departure of those forms owing to the

      expulsive pressure exerted by the surrounding air. Death, however,

      occurs not by haphazard but, when natural, owing to old age, and, when

      unnatural, to violence.

      But the reason for this and why all must die Democritus has by no

      means made clear. And yet, since evidently death occurs at one time of

      life and not at another, he should have said whether the cause is

      external or internal. Neither does he assign the cause of the

      beginning of respiration, nor say whether it is internal or

      external. Indeed, it is not the case that the external mind

      superintends the reinforcement; rather the origin of breathing and

      of the respiratory motion must be within: it is not due to pressure

      from around. It is absurd also that what surrounds should compress and

      at the same time by entering dilate. This then is practically his

      theory, and how he puts it.

      But if we must consider that our previous account is true, and

      that respiration does not occur in every animal, we must deem that

      this explains death not universally, but only in respiring animals.

      Yet neither is it a good account of these even, as may clearly be seen

      from the facts and phenomena of which we all have experience. For in

      hot weather we grow warmer, and, having more need of respiration, we

      always breathe faster. But, when the air around is cold and

      contracts and solidifies the body, retardation of the breathing

     
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