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      like an insect, has a body that admits of being curled up, its

      vertebrae being cartilaginous and easily bent. The faculty in question

      belongs then to serpents simply as a necessary consequence of this

      character of their vertebrae; but at the same time it has a final

      cause, for it enables them to guard against attacks from behind. For

      their body, owing to its length and the absence of feet, is ill-suited

      for turning round and protecting the hinder parts; and merely to

      lift the head, without the power of turning it round, would be of no

      use whatsoever.

      The animals with which we are dealing have, moreover, a part which

      corresponds to the breast; but neither here nor elsewhere in their

      body have they any mammae, as neither has any bird or fish. This is

      a consequence of their having no milk; for a mamma is a receptacle for

      milk and, as it were, a vessel to contain it. This absence of milk

      is not peculiar to these animals, but is common to all such as are not

      internally viviparous. For all such produce eggs, and the nutriment

      which in Vivipara has the character of milk is in them engendered in

      the egg. Of all this, however, a clearer account will be given in

      the treatise on Generation. As to the mode in which the legs bend, a

      general account, in which all animals are considered, has already been

      given in the dissertation on Progression. These animals also have a

      tail, larger in some of them, smaller in others, and the reason for

      this has been stated in general terms in an earlier passage.

      Of all oviparous animals that live on land there is none so lean

      as the Chamaeleon. For there is none that has so little blood. The

      explanation of this is to be found in the psychical temperament of the

      creature. For it is of a timid nature, as the frequent changes it

      undergoes in its outward aspect testify. But fear is a

      refrigeration, and results from deficiency of natural heat and

      scantiness of blood. We have now done with such sanguineous animals as

      are quadrupedous and also such as are apodous, and have stated with

      sufficient completeness what external parts they possess, and for what

      reason they have them.

      12

      The differences of birds compared one with another are differences

      of magnitude, and of the greater or smaller development of parts. Thus

      some have long legs, others short legs; some have a broad tongue,

      others a narrow tongue; and so on with the other parts. There are

      few of their parts that differ save in size, taking birds by

      themselves. But when birds are compared with other animals the parts

      present differences of form also. For in some animals these are hairy,

      in others scaly, and in others have scale-like plates, while birds are

      feathered.

      Birds, then, are feathered, and this is a character common to them

      all and peculiar to them. Their feathers, too, are split and

      distinct in kind from the undivided feathers of insects; for the

      bird's feather is barbed, these are not; the bird's feather has a

      shaft, these have none. A second strange peculiarity which

      distinguishes birds from all other animals is their beak. For as in

      elephants the nostril serves in place of hands, and as in some insects

      the tongue serves in place of mouth, so in birds there is a beak,

      which, being bony, serves in place of teeth and lips. Their organs

      of sense have already been considered.

      All birds have a neck extending from the body; and the purpose of

      this neck is the same as in such other animals as have one. This

      neck in some birds is long, in others short; its length, as a

      general rule, being pretty nearly determined by that of the legs.

      For long-legged birds have a long neck, short-legged birds a short

      one, to which rule, however, the web-footed birds form an exception.

      For to a bird perched up on long legs a short neck would be of no

      use whatsoever in collecting food from the ground; and equally useless

      would be a long neck, if the legs were short. Such birds, again, as

      are carnivorous would find length in this part interfere greatly

      with their habits of life. For a long neck is weak, and it is on their

      superior strength that carnivorous birds depend for their subsistence.

      No bird, therefore, that has talons ever has an elongated neck. In

      web-footed birds, however, and in those other birds belonging to the

      same class, whose toes though actually separate have flat marginal

      lobes, the neck is elongated, so as to be suitable for collecting food

      from the water; while the legs are short, so as to serve in

      swimming. The beaks of birds, as their feet, vary with their modes

      of life. For in some the beak is straight, in others crooked;

      straight, in those who use it merely for eating; crooked, in those

      that live on raw flesh. For a crooked beak is an advantage in

      fighting; and these birds must, of course, get their food from the

      bodies of other animals, and in most cases by violence. In such birds,

      again, as live in marshes and are herbivorous the beak is broad and

      flat, this form being best suited for digging and cropping, and for

      pulling up plants. In some of these marsh birds, however, the beak

      is elongated, as too is the neck, the reason for this being that the

      bird get its food from some depth below the surface. For most birds of

      this kind, and most of those whose feet are webbed, either in their

      entirety or each part separately, live by preying on some of the

      smaller animals that are to be found in water, and use these parts for

      their capture, the neck acting as a fishing-rod, and the beak

      representing the line and hook.

      The upper and under sides of the body, that is of what in quadrupeds

      is called the trunk, present in birds one unbroken surface, and they

      have no arms or forelegs attached to it, but in their stead wings,

      which are a distinctive peculiarity of these animals; and, as these

      wings are substitutes for arms, their terminal segments lie on the

      back in the place of a shoulder-blade.

      The legs are two in number, as in man; not however, as in man,

      bent outwards, but bent inwards like the legs of a quadruped. The

      wings are bent like the forelegs of a quadruped, having their

      convexity turned outwards. That the feet should be two in number is

      a matter of necessity. For a bird is essentially a sanguineous animal,

      and at the same time essentially a winged animal; and no sanguineous

      animal has more than four points for motion In birds, then, as in

      those other sanguineous animals that live and move upon the ground,

      the limbs attached to the trunk are four in number. But, while in

      all the rest these four limbs consist of a pair of arms and a pair

      of legs, or of four legs as in quadrupeds, in birds the arms or

      forelegs are replaced by a pair of wings, and this is their

      distinctive character. For it is of the essence of a bird that it

      shall be able to fly; and it is by the extension of wings that this is

      made possible. Of all arrangements, then, the only possible, and so

      the necessary, one is that birds shall have two feet; for this with

    &n
    bsp; the wings will give them four points for motion. The breast in all

      birds is sharp-edged, and fleshy. The sharp edge is to minister to

      flight, for broad surfaces move with considerable difficulty, owing to

      the large quantity of air which they have to displace; while the

      fleshy character acts as a protection, for the breast, owing to its

      form, would be weak, were it not amply covered.

      Below the breast lies the belly, extending, as in quadrupeds and

      in man, to the vent and to the place where the legs are jointed to the

      trunk.

      Such, then, are the parts which lie between the wings and the

      legs. Birds like all other animals, whether produced viviparously or

      from eggs, have an umbilicus during their development, but, when the

      bird has attained to fuller growth, no signs of this remain visible.

      The cause of this is plainly to be seen during the process of

      development; for in birds the umbilical cord unites with the

      intestine, and is not a portion of the vascular system, as is the case

      in viviparous animals.

      Some birds, again, are well adapted for flight, their wings being

      large and strong. Such, for instance, are those that have talons and

      live on flesh. For their mode of life renders the power of flight a

      necessity, and it is on this account that their feathers are so

      abundant and their wings so large. Besides these, however, there are

      also other genera of birds that can fly well; all those, namely,

      that depend on speed for security, or that are of migratory habits. On

      the other hand, some kinds of birds have heavy bodies and are not

      constructed for flight. These are birds that are frugivorous and

      live on the ground, or that are able to swim and get their living in

      watery places. In those that have talons the body, without the

      wings, is small; for the nutriment is consumed in the production of

      these wings, and of the weapons and defensive appliances; whereas in

      birds that are not made for flight the contrary obtains, and the

      body is bulky and so of heavy weight. In some of these heavy-bodied

      birds the legs are furnished with what are called spurs, which replace

      the wings as a means of defence. Spurs and talons never co-exist in

      the same bird. For nature never makes anything superfluous; and if a

      bird can fly, and has talons, it has no use for spurs; for these are

      weapons for fighting on the ground, and on this account are an

      appanage of certain heavy-bodied birds. These latter, again, would

      find the possession of talons not only useless but actually injurious;

      for the claws would stick into the ground and interfere with

      progression. This is the reason why all birds with talons walk so

      badly, and why they never settle upon rocks. For the character of

      their claws is ill-suited for either action.

      All this is the necessary consequence of the process of development.

      For the earthy matter in the body issuing from it is converted into

      parts that are useful as weapons. That which flows upwards gives

      hardness or size to the beak; and, should any flow downwards, it

      either forms spurs upon the legs or gives size and strength to the

      claws upon the feet. But it does not at one and the same time

      produce both these results, one in the legs, the other in the claws;

      for such a dispersion of this residual matter would destroy all its

      efficiency. In other birds this earthy residue furnishes the legs with

      the material for their elongation; or sometimes, in place of this,

      fills up the interspaces between the toes. Thus it is simply a

      matter of necessity, that such birds as swim shall either be

      actually web-footed, or shall have a kind of broad blade-like margin

      running along the whole length of each distinct toe. The forms,

      then, of these feet are simply the necessary results of the causes

      that have been mentioned. Yet at the same time they are intended for

      the animal's advantage. For they are in harmony with the mode of

      life of these birds, who, living on the water, where their wings are

      useless, require that their feet shall be such as to serve in

      swimming. For these feet are so developed as to resemble the oars of a

      boat, or the fins of a fish; and the destruction of the foot-web has

      the same effect as the destruction of the fins; that is to say, it

      puts an end to all power of swimming.

      In some birds the legs are very long, the cause of this being that

      they inhabit marshes. I say the cause, because nature makes the organs

      for the function, and not the function for the organs. It is, then,

      because these birds are not meant for swimming that their feet are

      without webs, and it is because they live on ground that gives way

      under the foot that their legs and toes are elongated, and that

      these latter in most of them have an extra number of joints. Again,

      though all birds have the same material composition, they are not

      all made for flight; and in these, therefore, the nutriment that

      should go to their tail-feathers is spent on the legs and used to

      increase their size. This is the reason why these birds when they

      fly make use of their legs as a tail, stretching them out behind,

      and so rendering them serviceable, whereas in any other position

      they would be simply an impediment.

      In other birds, where the legs are short, these are held close

      against the belly during flight. In some cases this is merely to

      keep the feet out of the way, but in birds that have talons the

      position has a further purpose, being the one best suited for

      rapine. Birds that have a long and a thick neck keep it stretched

      out during flight; but those whose neck though long is slender fly

      with it coiled up. For in this position it is protected, and less

      likely to get broken, should the bird fly against any obstacle.

      In all birds there is an ischium, but so placed and of such length

      that it would scarcely be taken for an ischium, but rather for a

      second thigh-bone; for it extends as far as to the middle of the

      belly. The reason for this is that the bird is a biped, and yet is

      unable to stand erect. For if its ischium extended but a short way

      from the fundament, and then immediately came the leg, as is the

      case in man and in quadrupeds, the bird would be unable to stand up at

      all. For while man stands erect, and while quadrupeds have their heavy

      bodies propped up in front by the forelegs, birds can neither stand

      erect owing to their dwarf-like shape, nor have anterior legs to

      prop them up, these legs being replaced by wings. As a remedy for this

      Nature has given them a long ischium, and brought it to the centre

      of the body, fixing it firmly; and she has placed the legs under

      this central point, that the weight on either side may be equally

      balanced, and standing or progression rendered possible. Such then

      is the reason why a bird, though it is a biped, does not stand

      erect. Why its legs are destitute of flesh has also already been

      stated; for the reasons are the same as in the case of quadrupeds.

      In all birds alike, whether web-footed or not, the number of toes in

      each foot is four. For the Liby
    an ostrich may be disregarded for the

      present, and its cloven hoof and other discrepancies of structure as

      compared with the tribe of birds will be considered further on. Of

      these four toes three are in front, while the fourth points

      backward, serving, as a heel, to give steadiness. In the long-legged

      birds this fourth toe is much shorter than the others, as is the

      case with the Crex, but the number of their toes is not increased. The

      arrangement of the toes is such as has been described in all birds

      with the exception of the wryneck. Here only two of the toes are in

      front, the other two behind; and the reason for this is that the

      body of the wryneck is not inclined forward so much as that of other

      birds. All birds have testicles; but they are inside the body. The

      reason for this will be given in the treatise On the Generation of

      Animals.

      13

      Thus then are fashioned the parts of birds. But in fishes a still

      further stunting has occurred in the external parts. For here, for

      reasons already given, there are neither legs nor hands nor wings, the

      whole body from head to tail presenting one unbroken surface. This

      tail differs in different fishes, in some approximating in character

      to the fins, while in others, namely in some of the flat kinds, it

      is spinous and elongated, because the material which should have

      gone to the tail has been diverted thence and used to increase the

      breadth of the body. Such, for instance, is the case with the

      Torpedos, the Trygons, and whatever other Selachia there may be of

      like nature. In such fishes, then, the tail is spinous and long; while

      in some others it is short and fleshy, for the same reason which makes

      it spinous and long in the Torpedo. For to be short and fleshy comes

      to the same thing as to be long and less amply furnished with flesh.

      What has occurred in the Fishing-frog is the reverse of what has

      occurred in the other instances just given. For here the anterior

      and broad part of the body is not of a fleshy character, and so all

      the fleshy substance which has been thence diverted has been placed by

     
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